Everything about Cephalochordata totally explained
The
lancelets (subphylum
Cephalochordata, traditionally known as
amphioxus) are a group of primitive
chordates. They are usually found buried in sand in shallow parts of
temperate or
tropical seas. In
Asia, they're harvested commercially for
food for
humans and
domesticated animals. They are an important object of study in
zoology as they provide indications about the origins of the
vertebrates.
Physical features
Lancelets grow up to about five
centimetres long, reaching eight centimetres at the longest. In common with vertebrates, lancelets have a nerve cord running along the back,
pharyngeal slits and a
tail that runs past the
anus. Also like vertebrates, the
muscles are arranged in blocks called
myomeres. Unlike vertebrates, the dorsal nerve cord isn't protected by
bone but by a simpler
notochord made up of a
cylinder of
cells that are closely-packed to form a toughened rod. The lancelet notochord, unlike the vertebrate
spine, extends into the head. This gives the subphylum its name (
cephalo- meaning 'relating to the head'). Lancelets also have oral
cirri, thin
tentacle-like strands that hang in front of the mouth and act as sensory devices and as a filter for the water passing into the body. The water exits the body via the
atriopore.
- brain-like blister
- notochord
- dorsal nerve cord
- post-anal tail
- anus
- food canal
- circulatory system
- atriopore
- overpharynx lacuna
- gill slit
- pharynx
- vestibule
- oral cirri
- mouth opening
- gonads (ovary/testicle)
- light sensor
- nerves
- metapleural fold
- hepatic caecum (liver-like sack)
Taxonomy
Cephalochordata is traditionally seen as a sister
subphylum to the
vertebrates, with whom they're grouped together as a sister group (sometimes called Notochordata) to the simpler still
Urochordata. But newer research suggests this may not be the case. Urochordates may be the sister group of the vertebrates, while Cephalochordata may be the most
basal subphylum of the chordates. The
asymmetric nature of juveniles is unique to the cephalochordates and indicates (as do certain other features, including the seriated
gonads) that lancelets are, at least, more derived than would be expected of a "living fossil" representative of basal chordates.
The following are the species recognised by
ITIS. Other sources (see for instance Tudge) show that there might be up to thirty species.
Family Asymmetronidae
- Genus Asymmetron
- Asymmetron lucayanum
- Asymmetron maldivense
- Genus Epigonichthys
Family Branchiostomidae
- Genus Branchiostoma
- Branchiostoma belcheri
- Branchiostoma californiense
- Branchiostoma capense
- Branchiostoma caribaeum
- Branchiostoma floridae
- Branchiostoma lanceolatum
- Branchiostoma valdiviae
- Branchiostoma virginiae
In music
Folk musician and marine biologist Sam Hinton performed a song attributed to Dr. Sewell H. Hopkins about Amphioxus entitled It's a Long Way from Amphioxus. Sung to the tune of "It's a Long Way to Tipperary," the ditty references now-disputed notions about the place of cephalochordates in the chordate evolutionary timeline:
It's a long way from Amphioxus,
It's a long way to us,
It's a long way from Amphioxus
To the meanest human cuss.
It's good-bye, fins and gill-slits,
Welcome, lungs and hair!
It's a long, long way from Amphioxus,
But we all came from there.Further Information
Get more info on 'Cephalochordata'.
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